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COCOA: Verbundprojekt COCOA - nutrient cocktail in the coastal zone

01.01.2014 - 31.12.2017
Project coordinated by:
Aarhus University
Project manager (IOW):
Prof. Dr. Maren Voß

COCOA will investigate physical, biogeochemical and biological processes along the coastal zone in the water column and in sediments in a combined and coordinated fashion to improve our understanding of the interaction of these processes on the removal of nutrients along the land-sea interface. Our goal is to achieve true inter-disciplinarity through close collaborations, building our hypotheses together and incorporating the new knowledge generated into state-of-the-art ecosystem models. This coordinated action will allow us to synthesise the results on the entire ecosystem scale of the Baltic and to improve current decision support tools used for the implementation of the HELCOM BSAP, EU WFD and MSFD.


  • Saniewska, D., M. Bełdowska, E. Szymczak, K. Kuliński, J. Bełdowski, M. Voss, D. Pryputniewicz-Flis and D. Burska (2022). Processes affecting the transformation of mercury in the coastal zone in the vicinity of two river mouths in the southern Baltic Sea. Mar. Chem. 238: 104065, doi: 10.1016/j.marchem.2021.104065
  • Kache, S., I. Bartl, J. Wäge-Recchioni and M. Voss (2021). Influence of organic particle addition on nitrification rates and ammonium oxidiser abundances in Baltic seawater. Mar. Ecol. Prog. Ser. 674: 59-72, doi: 10.3354/meps13797
  • Zilius, M., I. Vybernaite-Lubiene, D. Vaiciute, D. Overlingė, E. Grinienė, A. Zaiko, S. Bonaglia, I. Liskow, M. Voss, A. Andersson, S. Brugel, T. Politi and P. A. Bukaveckas (2021). Spatiotemporal patterns of N2 fixation in coastal waters derived from rate measurements and remote sensing. Biogeosciences 18: 1857-1871, doi: 10.5194/bg-18-1857-2021
  • Broman, E., M. Zilius, A. Samuiloviene, I. Vybernaite-Lubiene, T. Politi, I. Klawonn, M. Voss, F. J. A. Nascimento and S. Bonaglia (2021). Active DNRA and denitrification in oxic hypereutrophic waters. Water Res. 194: 116954, doi: 10.1016/j.watres.2021.116954
  • Voss, M., E. Asmala, I. Bartl, J. Carstensen, D. J. Conley, J. W. Dippner, C. Humborg, K. Lukkari, J. Petkuviene, H. Reader, C. Stedmon, I. Vybernaite-Lubiene, N. Wannicke and M. Zilius (2021). Origin and fate of dissolved organic matter in four shallow Baltic Sea estuaries. Biogeochemistry 154: 385-403, doi: 10.1007/s10533-020-00703-5
  • Dippner, J. W., I. Bartl, E. Chrysagi, P. Holtermann, A. Kremp, F. Thoms and M. Voss (2019). Lagrangian residence time in the Bay of Gdańsk, Baltic Sea. Front. Mar. Sci. 6: 725, doi: 10.3389/fmars.2019.00725
  • Bartl, I., D. Hellemann, C. Rabouille, K. Schulz, P. Tallberg, S. Hietanen and M. Voss (2019). Particulate organic matter controls benthic microbial N retention and N removal in contrasting estuaries of the Baltic Sea. Biogeosciences 16: 3543-3564, doi: 10.5194/bg-16-3543-2019
  • Carstensen, J., D. J. Conley, E. Almroth-Rosell, E. Asmala, E. Bonsdorff, V. Fleming-Lehtinen, B. G. Gustafsson, C. Gustafsson, A.-S. Heiskanen, U. Janas, A. Norkko, C. Slomp, A. Villnas, M. Voss and M. Zilius (2020). Factors regulating the coastal nutrient filter in the Baltic Sea. Ambio 49: 1194-1210, doi: 10.1007/s13280-019-01282-y
  • Asmala, E., J. Carstensen, D. J. Conley, C. P. Slomp, J. Stadmark and M. Voss (2019). A reply to the comment by Karlsson et al. Limnol. Oceanogr. 64: 1832-1833, doi: 10.1002/lno.11195
  • Reader, H. E., F. Thoms, M. Voss and C. A. Stedmon (2019). The influence of sediment-derived dissolved organic matter in the Vistula river estuary/Gulf of Gdansk. J. Geophys. Res. Biogeosciences 124: 115-126, doi: 10.1029/2018jg004658
  • Zilius, M., I. Vybernaite-Lubiene, D. Vaiciute, J. Petkuviene, P. Zemlys, I. Liskow, M. Voss, M. Bartoli and P. A. Bukaveckas (2018). The influence of cyanobacteria blooms on the attenuation of nitrogen throughputs in a Baltic coastal lagoon. Biogeochemistry 141: 143-165, doi: 10.1007/s10533-018-0508-0
  • Thoms, F., C. Burmeister, J. W. Dippner, M. Gogina, U. Janas, H. Kendzierska, I. Liskow and M. Voss (2018). Impact of macrofaunal communities on the coastal filter function in the Bay of Gdansk, Baltic Sea. Front. Mar. Sci. 5: 201, doi: 10.3389/fmars.2018.00201
  • Bartl, I., I. Liskow, K. Schulz, L. Umlauf and M. Voss (2018). River plume and bottom boundary layer - Hotspots for nitrification in a coastal bay? Estuar. Coast. Shelf Sci. 208: 70-82, doi: 10.1016/j.ecss.2018.04.023
  • Umbricht, J., J. W. Dippner, B. Fry, I. Kröncke, I. Liskow, P. Nehmer, F. Thoms and M. Voss (2018). Correction of the isotopic composition (δ13C and δ15N) of preserved Baltic and North Sea macrozoobenthos and their trophic interactions. Mar. Ecol. Prog. Ser. 595: 1-13, doi: 10.3354/meps12543
  • Hellemann, D., P. Tallberg, I. Bartl, M. Voss and S. Hietanen (2017). Denitrification in an oligotrophic estuary: a delayed sink for riverine nitrate. Mar. Ecol. Prog. Ser. 583: 63-80, doi: 10.3354/meps12359
  • Asmala, E., J. Carstensen, D. J. Conley, C. P. Slomp, J. Stadmark and M. Voss (2017). Efficiency of the coastal filter: Nitrogen and phosphorus removal in the Baltic Sea. Limnol. Oceanogr. 62: S222-S238, doi: 10.1002/lno.10644
  • Xenopoulos, M. A., J. A. Downing, M. D. Kumar, S. Menden-Deuer and M. Voss (2017). Headwaters to oceans: Ecological and biogeochemical contrasts across the aquatic continuum. Limnol. Oceanogr. 62: S3-S14, doi: 10.1002/lno.10721